Hernandez-Castillo, Genaro R. , Stockey, Ruth A. , Rothwell, Gar W. , Mapes, Gene .
A whole plant reconstruction of the Upper Pennsylvanian conifer Emporia lockardii (Emporiaceae).
THE fossil flora from the Hamilton Quarry in south eastern Kansas includes a diversity of conifers and conifer-like plants from which three species have been described (Emporia lockardii, Barthelia furcata, and Hanskerpia hamiltonensis). Emporia lockardii was originally described from permineralized ovulate cones attached to leafy branches. In this paper we expand the species concept to include growth architecture, lateral branch morphology, shoot anatomy, leaf form and diversity, cuticular anatomy, pollen cone structure, and pollen. This species is represented by eighty five compression/impression and permineralized specimens of vegetative shoots, pollen cones, and ovulate cones. Emporia lockardii has an orthotropic stem that produces lateral plagiotropic branches with simple and forked leaves, simple pollen cones, and compound ovulate cones. Stems have an endarch eustele with dense wood surrounding a septate pith. Leaves display age-dependent heterophylly, and are amphistomatic with two adaxial stomatal bands, papillate epidermal cells, and either two narrow, basal stomatal areas or short rows of stomata with numerous trichome bases abaxially. Pollen cones have helically arranged peltate microsporophylls and adaxial pollen sacs. Prepollen is monolete and monosaccate, and conforms to the sporae dispersae genus Potonieisporites Bharadwaj. Ovulate cones are compound with helically arranged, forked bracts that subtend bilateral dwarf shoots with numerous sterile scales and one to three narrow megasporophylls. Ovules are terminal, inverted, and somewhat flattened. This set of diagnostic characters clearly sets the genus Emporia apart from other walchian conifers, and provides evidence for a reevaluation of the family Emporiaceae.
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1 - Universidad Nacional Autonoma de Mexico, Departamento de Paleontología, Instituto de Geología, México City, 04510, México
2 - University of Alberta, Department of Biological Sciences, Biological Sciences Centre, Cw 405, Edmonton, Alberta, T6G 2E9, Canada
3 - Ohio University, Department of Environmental & Plant Biology, Porter Hall, Richland Avenue, Athens, Ohio, 45701-2979, USA
walchian conifer plant.
Presentation Type: Oral Paper:Papers for Sections
Date: Monday, July 31st, 2006
Time: 1:15 PM